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Owl box - could a red squirrel commandeer it?

I have a large lime tree in my garden which has had an owl box on it for years - no owls have ever nested in it although we have a good population of tawneys in the neighbourhood. Yesterday whilst out walking I noticed that my owl box had lots of small branches poking through the opening - very unusual I thought to myself. Whilst watching for the BGBW this morning I saw two red squirrels arguing over the peanut feeder and then later one of them at the very top of the lime tree looking as if it was collecting twigs. Never saw it reappear at the bottom of the tree...could it be preparing a drey in the nestbox? I know this is a bird forum but thought if you liked birds some of you may also have an interest in mammals....

thanks in advance for any insights